故事是这样的：小老鼠菲维尔在纽约安定下来后，听说西部有个没有猫的老鼠镇，便带着家人踏上火车，一路向西。 影片是典型的手绘西部动画，线条粗犷却细腻，火车穿越荒野的镜头透着一股老式冒险片的朴实劲。旅途中他结识了孤独的牛仔猫、会唱歌的草原犬鼠，还有总想使坏的响尾蛇团伙。路上他们遇到了会唱歌的沙漠蜥蜴、爱吹牛的野牛，还有一只总想偷火车奶酪的坏猫。音乐段落里的口琴和班卓琴很有味道，尤其是野牛合唱的那段“西部之路”，简单得就像篝火旁随口哼出来的调子。剧情没有说教，靠的就是伟福一次次笨拙但真诚的尝试——比如他为了穿过蛇洞，竟想出用仙人掌汁液润滑的方法。

Some time after the Mousekewitz's have settled in America, they find that they are still having problems with the threat of cats. That makes them eager to try another home out in the west, where they are promised that mice and cats live in peace. Unfortunately, the one making this claim is an oily con artist named Cat R. Waul who is intent on his own sinister plan. Unaware of this, the Mousekewitz's begin their journey west, while their true cat friend, Tiger, follows intent on following his girlfriend gone in the same direction.

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